No, the 270 bullet will NOT out penetrate a 480-500 gr., 45 cal. bullet in anything, flesh, mud, sand, green cheese or one eyed, one eared purple people eaters. It isn't I who misunderstand anything.
I have to ask, how is it you assume my front to rear shots and those I've witnessed "usually" did not take out any bone? Not all did however, just as many did shatter shoulder blades or wreck the pelvic, penetrated the ham front to rear or slid along the spine taking off tips of vertabrae as they passed...and those were were all cast bullets. Those cast bullets, none caused anywhere near the "massive bleeding" one sees from expanding bullets at higher velocities. Nor do the solids used on dangerous game. They have one purpose and one purpose only, to penetrate heavy muscle, bone and stay on course to reach vitals and kill the animal. The aren't to expand, rivet, bend or anything else, just penetrate. At 1100-1200 fps a cast bullet acts much the same which is why the importance, to me, of a large meplat.
With 270 class cartridges I also had and witnessed more than a few broadside shots. Those that were, penetration was complete, side to side, whether a bone or several bones, shoulder blade/ribs, were hit. My wife's 243 has done the same, every time she's taken that shot. Bullet fragments were sometimes found but never remains of the entire bullet.
Tell ya what, I've done these penetration "tests" in wet phone books and wet paper and baffle board boxes...and more than a few critters. Ain't no 270, '06, 280, 44 Mag, 357 Mag, 243 or 7mm Mag. gonna out penetrate long, heavy for caliber, cast bullets at moderate velocity whether that be in ballistic gelatin, wet books, wet or dry paper, baffle boxes or game.....or mud. It ain't gonna happen, period.